/ Dictionary / Index C columella: Meaning and Definition of Find definitions for: col•u•mel•la Pronunciation: (kol"yu-mel'u), [key] — pl. -mel•lae any of various small, columnlike structures of animals or plants; rod or axis. Mycol.a small central column of sterile tissue within the sporangium of certain fungi, liverworts, and mosses. a small bone in the ear of amphibians, reptiles, and birds. Random House Unabridged Dictionary, Copyright © 1997, by Random House, Inc., on Infoplease. See also: columella (Thesaurus) Columbus Day columelliform Related Content Daily Word Quiz: vertigo Analogy of the Day: Today’s Analogy Spelling Bee: Today’s Spelling Bee Frequently Misspelled Words Frequently Mispronounced Words Easily Confused Words Writing & Language