/ Dictionary / Index C complementary base: Meaning and Definition of Find definitions for: com'plemen'tary base' Pronunciation: [key] — Genetics. Genetics. either of the nucleotide bases linked by a hydrogen bond on opposite strands of DNA or double-stranded RNA: guanine is the complementary base of cytosine, and adenine is the complementary base of thymine in DNA and of uracil in RNA. Cf. base pair. Random House Unabridged Dictionary, Copyright © 1997, by Random House, Inc., on Infoplease. complementary angle complementary cells Related Content Daily Word Quiz: sumptuous Analogy of the Day: Today’s Analogy Spelling Bee: Today’s Spelling Bee Frequently Misspelled Words Frequently Mispronounced Words Easily Confused Words Writing & Language