Cnidaria

Introduction

CE5

Internal anatomy of Hydra, representative of the phylum Cnidaria

Cnidaria səlĕntəräˈtə [key], phylum of invertebrate animals comprising the sea anemones, corals, jellyfish, and hydroids. Cnidarians are radially symmetrical (see symmetry, biological). The mouth, located at the center of one end of the body, opens into a gastrovascular cavity, which is used for digestion and distribution of food; an anus is lacking. Cnidarians are further characterized by having a body wall composed of three layers: an outer epidermis, an inner gastrodermis, and a middle mesogloea. Tentacles encircle the mouth and are used in part for food capture. Specialized stinging structures, called nematocysts, are a characteristic of the phylum and are borne in the tentacles and often in other body parts. These contain a coiled fiber that can be extruded suddenly. Some nematocysts contain toxic substances and are defense mechanisms, while others are adhesive, helping to anchor the animal or to entangle prey.

Two body forms and two lifestyles are characteristic of the Cnidaria (see polyp and medusa). The sessile hydroid, or polyp, form is more or less cylindrical, attached to its substratum at its aboral (opposite the mouth) end, with the mouth and surrounding tentacles at the upper, oral, free end. Colonies of hydroids comprise several different types of individuals: some function in feeding, some in defense, and some in reproduction. The motile jellyfish, or medusoid form, is flattened, with the tentacles usually located at the body margin. The medusoid's convex aboral surface is oriented upward, and the concave oral surface is oriented downward.

With few exceptions, the cnidarians are marine. There are over 9,000 known living species; fossil records of cnidarians date back to the Ordovician era. Cnidarians are carnivorous, the major part of their diet consisting of crustaceans. Animals in this phylum have no specialized excretory or respiratory organs but do have a nervous system. Both sexual and asexual reproduction occur. There are three classes of cnidarians.

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